享保6年，将军吉宗下令设置了“意见箱”。藉由投递至意见箱的投书，新的奉行所诞生了。又名“虫奉行所”！为了保护人们免于遭受盘据江户的大虫所害，被选出的强者们大显神通。继承父亲的志向来到江户的菜鸟同心·仁兵卫！然而他还不知道虫奉行所令人惊讶的工作内容……
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