故事叙述在“东京大决战”结束十年之后，日本各地区已经成为了一个个的独立国家。而一名任职于自警团、成绩普通的女孩森友望未，某日心血来潮地踏上了横跨日本列岛诸国的旅程。
她沿途邂逅了响逢衣、御园千绫、小坂结季奈等人，这四个女孩将背负起守护和平的使命。而在旅途前方等待着他们的有魔法师、怪兽与巨大机器人!?由这四名少女所交织而成，新感觉的公路青春故事即将上演！
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